• BIOL 4200

    Networks in Biology
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    Fall 2026

    Why do some diseases become global pandemics, while others die out? What happens to a food web when a species goes extinct? What roles do different organisms have within a social network? How does the structure of brain networks relate to cognitive function? Networks are all around us; we just need to learn to see them. This course will introduce network analysis as a tool for answering fundamental and applied questions across biological disciplines. Coding experience is strongly recommended.

  • BIOL 4300

    The Origin of Species
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    Last Taught

    Fall 2026

    The process of speciation is fundamental to understanding life's diversity, with implications for how we study evolution, how we define species in nature, and the processes by which they form. In this course, we will explore models of adaptive and non-adaptive speciation and how we test and distinguish among these processes. We will explore examples from biological systems in nature, including syntheses of broad knowledge gained about the speciation process from studying each group of organisms.

  • BIOL 8010

    Integrative Biology Colloquium
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    Fall 2026

    A weekly conference in which students present reports covering various aspects of Integrative Biology. May be repeated for credit.

  • BIOL 8011

    Integrative Biology I
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    Last Taught

    Fall 2026

    Integrative Biology seeks to understand how genetic, cellular, and developmental processes give rise to organismal phenotypes and how changes in these processes over time contribute to the evolution of form and function. In this course, graduate students will learn principles of genetics, cell, developmental, and organismal biology and apply this knowledge towards answering fundamental questions in biology.

  • BIOL 8070

    Colloquium in Population Biology
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    Last Taught

    Fall 2026

    A weekly conference arranged around a current topic. May be repeated for credit. Prerequisite: Instructor permission.

  • BIOL 8999

    Non-Topical Research
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    Last Taught

    Fall 2026

    For master's thesis, taken under the supervision of a thesis director.

  • BIOL 9995

    Topical Research in Biology
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    Last Taught

    Fall 2026

    Independent research with a member of the Biology faculty in preparation for thesis or dissertation research.

  • BIOL 9998

    Non-Topical Research, Preparation for Doctoral Research
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    Last Taught

    Fall 2026

    For doctoral research, taken before a dissertation director has been selected.

  • BIOL 9999

    Non-Topical Research
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    Last Taught

    Fall 2026

    For doctoral dissertation, taken under the supervision of a dissertation director.